The mullets or grey mullets are a family (Mugilidae) of ray-finned fish found worldwide in coastal temperate and tropical waters, and some species in fresh water.
Mullets have served as an important source of food in Mediterranean Europe since Roman times.
The family includes about 78 species in 20 genera.
Mullets are distinguished by the presence of two separate dorsal fins, small triangular mouths, and the absence of a lateral line organ.
They feed on detritus, and most species have unusually muscular stomachs and a complex pharynx to help in digestion.
Classification and naming
thumb|right| Thick lips of a mullet thumb|right| Mullets in the Mediterranean Sea
Taxonomically, the family is currently treated as the sole member of the order Mugiliformes, but as Nelson says, "there has been much disagreement concerning the relationships" of this family.
The presence of fin spines clearly indicates membership in the superorder Acanthopterygii, and in the 1960s, they were classed as primitive perciforms,Gosline, W. A. (1961)
"The Perciform Caudal Skeleton"  Copeia 1961(3): pp.
265-270 while others have grouped them in Atheriniformes.
They are classified as an order, Mugiliformes, within the subseries Ovalentaria of the clade Percomorpha in the 5th Edition of Fishes of the World.
In North America, "mullet" by itself usually refers to Mugilidae.
In Europe, the word "mullet" is usually qualified, the "grey mullets" being Mugilidae and the "red mullets" or "surmullets" being Mullidae, notably members of the genus Mullus.
Outside Europe, the Mullidae are often called "goatfish".
Fish with common names including the word "mullet" may be a member of one family or the other, or even unrelated such as the freshwater white sucker (Catostomus commersonii).
However, recent taxonomic work has reorganised the family and the following genera make up the Mugilidae:
Agonostomus Bennett, 1832
Aldrichetta Whitley, 1945
Cestraeus Valenciennes, 1836
Chaenomugil Gill, 1863
Chelon Artedi, 1763
Crenimugil Schultz, 1946
Dajaus Valenciennes, 1836
Ellochelon Whitley, 1930
Gracilimugil Whitley, 1941
Joturus Poey, 1860
Minimugil Durand, Chen, Shen, Fu & Borsa, 2012
Mugil Linnaeus, 1758
Myxus  Günther, 1861
Neomyxus Steindachner, 1878
Neochelon Durand, Chen, Shen, Fu & Borsa 2012
Oedalechilus Fowler 1903
Osteomugil G. Luther, 1982
Parachelon Durand, Chen, Shen, Fu & Borsa 2012
Paramugil Ghasemzadeh, Ivantsoff & Aarn 2004
Planiliza Whitley, 1945
Plicomugil Schultz, 1953
Pseudomyxus Durand, Chen, Shen, Fu & Borsa 2012
Rhinomugil Gill, 1863
Sicamugil Fowler, 1939
Squalomugil Ogilby, 1908
Trachystoma Ogilby, 1888
Behaviour
A common noticeable behaviour in mullet is the tendency to leap out of the water.
There are two distinguishable types of leaps: a straight, clean slice out of the water to escape predators and a slower, lower jump while turning to its side that results in a larger, more distinguishable, splash.
The reasons for this lower jump are disputed, but have been hypothesised to be in order to gain oxygen rich air for gas exchange in a small organ above the pharynx.
Development
The ontogeny of mugilid larvae has been well studied, with the larval development of Mugil cephalus in particular being studied intensively due to its wide range of distribution and interest to aquaculture.
The previously understudied osteological development of Mugil cephalus was investigated in a 2021 study, with four embryonic and six larval developmental steps being described in aquaculture-reared and wild-caught specimens.
These descriptions provided clarification of questionable characters of adult mullets and revealed informative details with potential implications for phylogenetic hypotheses, as well as providing an overdue basis of comparison for aquaculture-reared mullets to enable recognition of malformations.
Timeline
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bar:eratop  from: -65.5   till:  -23.03    color:paleogene  text:[[Paleogene]]           from: -23.03    till: -2.588    color:neogene    text:[[Neogene]]  from: -2.588    till: 0   color:quaternary    text:[[Quaternary|Q.]]
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color:oligocene bar:NAM1  from:	-33.9	till:	0	text:	[[Mugil]]  color:miocene bar:NAM2  from:	-23.03	till:	0	text:	[[Liza (fish)|Liza]]
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bar:period  from: -65.5   till:  -55.8    color:paleocene  text:[[Paleocene]]  from: -55.8   till:  -33.9    color:eocene  text:[[Eocene]]   from: -33.9   till:  -23.03    color:oligocene  text:[[Oligocene]]              from: -23.03    till: -5.332    color:miocene    text:[[Miocene]]  from: -5.332    till: -2.588    color:pliocene    text:[[Plio.]]  from: -2.588    till: -0.0117   color:pleistocene    text:[[Pleist.]]  from: -0.0117    till: 0    color:holocene    text:[[Holocene|H.]]
bar:era  from: -65.5   till:  -23.03    color:paleogene  text:[[Paleogene]]           from: -23.03    till: -2.588    color:neogene    text:[[Neogene]]  from: -2.588    till: 0   color:quaternary    text:[[Quaternary|Q.]]
References
Further references
J.S. Nelson, Fishes of the World. .
External links
Video: Mullet Dursey Sound May 2010, Beara, West Cork, Ireland
